Publications
- Royer, A.M., A. Lacina, and C.A.F. Howey. In press. Male dispersal drives gene flow in timber rattlesnakes (Crotalus horridus). Ichthyology and Herpetology.
- MacLeod, K.J., T. Langkilde, J.J. Hepner, C.A.F. Howey, K. Sprayberry, C. Tylan, and M.J. Sheriff. 2021. Compensating for a stressful pregnancy? Glucocorticoid treatment during gravidity reduces metabolic rate in female fence lizards post-partuition. Hormones and Behavior 136.
- Howey, C.A.F., and E. Snyder. 2020. Substrate type affects scent-trailing behavior of newborn timber rattlesnakes (Crotalus horridus). Copeia 108:772-777.
- Herr, M., J. Avery, T. Langkilde, and C.A.F. Howey. 2020. Trade-off between thermal quality and predation risk at timber rattlesnake gestation sites. Journal of Herpetology 54:196-205.
- Hromada, S.J., C.A.F. Howey, M.B. Dickinson, R.W. Perry, W.M. Roosenburg, C.M. Gienger. 2018. Response of reptile and amphibian communities to the reintroduction of fire in an oak/hickory forest. Forest Ecology and Management 428(2018):1-13.
- Howey, C.A.F., and Z. Maisch. 2017. Defense of a female hotspot by a male timber rattlesnake. Herpetological Review 48(1):16-19.
- Howey, C.A.F., M.B. Dickinson, and W.M. Roosenburg. 2017. The effects of a landscape disturbance on the habitat use and behavior of the black racer. Copeia 104(4):853-863.
- Hoekstra, A.A., and C.A.F. Howey. 2016. Crotalus horridus (Timber Rattlesnake): Reproductive/Combat Behavior. Herpetological Review 47(2):309.
- Howey, C.A.F., and W.M. Roosenburg. 2013. Effects of prescribed fire on eastern box turtles (Terrapene carolina carolina). Northeastern Naturalist 20(3):493-497.
- Howey, C.A.F., and S.A. Dinkelacker. 2013. Characteristics of a historically harvested alligator snapping turtle (Macrochelys temminckii) population. Copeia 2013(1):58-63.
- Howey, C.A.F., and W.M. Roosenburg. 2012. The effects of prescribed burning on the black kingsnake: going beyond the demographic data. In Proceedings of the 4th Fire in Eastern Oak Forests Conference. Editors: D. Dey, M. Stambaugh, S. Clark, and C. Schweitzer. Springfield, MO. May 17-19, 2011.
- Howey, C.A.F., and S.A. Dinkelacker. 2009. Habitat selection of the alligator snapping turtle (Macrochelys temminckii) in Arkansas. Journal of Herpetology 43(4):589-596.
- Howey, C.A.F., and S.A. Dinkelacker. 2007. Macrochelys temminckii (Alligator Snapping Turtle). Terrestrial Burrow Use. Herpetological Review 38(3):333-334.
- Howey, C.A.F., and S.A. Dinkelacker. 2007. New distributional records for reptiles in central Arkansas. Herpetological Review 38(2):237-238.
Presentations – Public Outreach
- Howey, C.A.F. The World of the Timber Rattlesnake. Lackawanna State Park. 11 August 2023.
- Howey, C.A.F. The World of the Timber Rattlesnake. Lacawac Sanctuary. 30 July 2023.
- Howey, C.A.F. The World of the Timber Rattlesnake: History, Ecology, and Conservation of America’s Snake. Lackawanna Audubon Society. 16 February 2020.
- Howey, C.A.F. The World of the Timber Rattlesnake. PPL Northeast Safety Forum. 15 May 2019.
- Howey, C.A.F. The World of Timber Rattlesnakes! Lackawanna College Environmental Education Center. 10 April 2019.
- Howey, C.A.F. The World of Timber Rattlesnakes! Lackawanna State Park. 12 March 2019.
- Howey, C.A.F. The World of Timber Rattlesnakes. The 2017 Allegany Nature Pilgrimage. 2-4 June 2017.
- Howey, C.A.F. Reptiles and Amphibians of Pennsylvania. The 2017 Wildlife Society Northeast Student Conclave. 8 April 2017.
Presentations – Invited
- Howey, C.A.F. History and ecology of timber rattlesnakes in an ever-changing world. Hampton University. 8 October 2020.
- Howey, C.A.F. History and ecology of timber rattlesnakes in an ever-changing world. Bighamton University. 7 December 2019.
- Howey, C.A.F. Restoration of timber rattlesnake rookeries: efficacy of daylighting management. Timber Rattlesnake Conservation Council. 29 September 2017.
- Howey, C.A.F. Disturbing rattlesnakes: how landscape disturbance affect timber rattlesnake populations. University of Maine. 3 April 2017.
- Howey, C.A.F. Ecological effects of landscape disturbance on timber rattlesnakes. USFS Warren Forestry Science Lab Research Seminar. 3 March 2017.
- Howey, C.A.F., W.M. Roosenburg, and M.B. Dickinson. Ecological ramifications of prescribed fire on the black racer (Coluber constrictor). The Wildlife Society Annual Meeting. 5-10 October 2013.
- Howey, C.A.F., W.M. Roosenburg, and M.B. Dickinson. Habitat use, movement rates, and energetic expenditures of black racers (Coluber constrictor) in burned and unburned landscapes. Austin Peay University. 11 February 2013.
- Howey, C.A.F., W.M. Roosenburg, and M.B. Dickinson. Fire effects on reptiles, their landscape, and habitat use. USFS Delaware Forestry Science Lab Research Seminar. 16 October 2012.
- Howey, C.A.F., W.M. Roosenburg, and M.B. Dickinson. Fire effects on reptiles and their habitat. 4th Annual Kentucky Prescribed Fire Council Meeting. 17-18 September 2012.
- Howey, C.A.F., W.M. Roosenburg., and M.B. Dickinson. Fire effects on snake habitat in Kentucky forests. Fire and Forest Management Impacts on Timber Rattlesnakes and Their Surrogates Council. 8 March 2012.
Presentations – Conference
* represents undergraduate coauthors. The first author is always the individual that delivered the presentation unless otherwise noted.
- Howey, C.A.F., and A.M. Royer. Male spatial use drives gene flow in timber rattlesnakes (Crotalus horridus). Joint Meeting of Ichthyologists and Herpetologists. 12-16 July 2023. (Oral Presentation)
- Howey, C.A.F. Thermoregulation and foraging behavior of timber rattlesnakes (Crotalus horridus) in a disturbed landscape. Society for Integrative and Comparative Biology. 3-7 January 2020. (Oral Presentation)
- Christiano, B.C.*, and C.A.F. Howey. Timber rattlesnakes (Crotalus horridus) that move more often maintain higher baseline corticosterone levels. Society for Integrative and Comparative Biology. 3-7 January 2020. (Oral Presentation)
- D’Allessandro, M.N.*, and C.A.F. Howey. Metabolic rates of city anoles. Society for Integrative and Comparative Biology. 3-7 January 2020. (Poster Presentation)
- Diorio, R.D.*, and C.A.F. Howey. The persistence of resistance to timber rattlesnake venom in small mammals. Society for Integrative and Comparative Biology. 3-7 January 2020. (Poster Presentation)
- Kenny, E.*, and C.A.F. Howey. Ambient light at night and effects on the immune response of anoles. Society for Integrative and Comparative Biology. 3-7 January 2020. (Poster Presentation)
- McGrath, M.R.*, and C.A.F. Howey. Ambient light at night (ALAN) and the stress response of green anoles. Society for Integrative and Comparative Biology. 3-7 January 2020. (Poster Presentation)
- Howey, C.A.F. Restoration of timber rattlesnake gestation sites: efficacy of daylighting management. Society for Integrative and Comparative Biology. 3-7 January 2019. (Oral Presentation)
- Christiano, B.C.*, and C.A.F. Howey. Do timber rattlesnakes with larger home ranges maintain higher corticosterone levels? Society for Integrative and Comparative Biology. 3-7 January 2019. (Poster Presentation)
- Diorio, R.D.*, and C.A.F. Howey. Does substrate type affect scent-trailing behavior of adult and newborn timber rattlesnakes? Society for Integrative and Comparative Biology. 3-7 January 2019. (Poster Presentation).
- Howey, C.A.F., and E. Snyder*. Substrate type affects scent-trailing behavior of timber rattlesnakes. Joint Meeting of Ichthyologists and Herpetologists. 11-15 July 2018. (Oral Presentation)
- Howey, C.A.F. Restoration of timber rattlesnake gestation sites: efficacy of daylighting management. Society for Integrative and Comparative Biology. 3-7 January 2018. (Oral Presentation)
- Novak, R.A.*, and C.A.F. Howey. A comparison of macroinvertebrate communities among vernal pools with varying fire histories. Society for Integrative and Comparative Biology. 3-7 January 2018. (Oral Presentation).
- Howey, C.A.F. Restoration of timber rattlesnake rookeries: efficacy of daylighting management. Joint Meeting of Ichthyologists and Herpetologists. 12-16 July 2017. (Oral Presentation)
- Hromada, S.J., C.A.F. Howey, and C.M. Gienger. Response of reptile and amphibian communities to prescribed fire in oak/hickory forests.Joint Meeting of Ichthyologists and Herpetologists. 12-16 July 2017. (Oral Presentation)
- Schneider, A.E., and C.A.F. Howey. Identification of individual timber rattlesnakes using camera trap data. Joint Meeting of Ichthyologists and Herpetologists. 12-16 July 2017. (Poster Presentation)
- Snyder, S.*, W. Sutton, and C.A.F. Howey. Differentiation in basking habitat between yellow and black morph timber rattlesnakes. Joint Meeting of Ichthyologists and Herpetologists. 12-16 July 2017. (Oral presentation)
- Howey, C.A.F., and W.M. Roosenburg. Effect of temperature on snake locomotion and the interpretation of thermal performance curves. Society for Integrative and Comparative Biology. 4-8 January 2017. (Oral Presentation)
- Mead, M.S., C.A.F. Howey, and T. Langkilde. Impacts of pH and UVB on stress and development rates of wood frog tadpoles: implications with regard to prescribed fire. Society for the Integrative and Comparative Biology. 4-8 January 2017. (Oral Presentation)
- Howey, C.A.F., M.S. Mead*, M.W. Herr*, and T. Langkilde. Thermoregulation and predation risk trade-offs at timber rattlesnake rookeries. Joint Meeting of Ichthyologists and Herpetologists. 6-10 July 2016. (Oral Presentation)
- Howey, C.A.F., W.M. Roosenburg, and M.B. Dickinson. Ecological effects of a disturbance event on the black racer (Coluber constrictor) habitat use, thermoregulation, and energetic expenditures. Ecological Society of America Annual Meeting. 9-14 August 2015. (Oral Presentation)
- Howey, C.A.F., W.M. Roosenburg, and M.B. Dickinson. Thermal preference, performance, and kinematics of the black racer. Society for the Study of Amphibians and Reptiles Annual Meeting. 30 July – 2 August 2015. (Poster Presentation)
- Howey, C.A.F., W.M. Roosenburg, and M.B. Dickinson. Thermoregulation and energy expenditures of black racers (Coluber constrictor) in thermally contrasting, burned and unburned landscapes. Joint Meeting of Ichthyologists and Herpetologists. 30 July – 3 August 2014. (Oral Presentation)
- Howey, C.A.F., W.M. Roosenburg, and M.B. Dickinson. Habitat use, energy expenditures, and body condition of black racers (Coluber constrictor) in burned and unburned landscapes. Joint Meeting of Ichthyologists and Herpetologists. 10-15 July 2013. (Oral Presentation)
- Howey, C.A.F., W.M. Roosenburg, and M.B. Dickinson. The effects of prescribed burning on the landscape and reptile abundance. Joint Meeting of Ichthyologists and Herpetologists. 10-15 July 2013. (Poster Presentation)
- Howey, C.A.F., W.M. Roosenburg, and M.B. Dickinson. The effects of prescribed burning on the landscape and reptile abundance. Society for Integrative and Comparative Biology. 3-7 January 2013. (Oral Presentation)
- Howey, C.A.F., W.M. Roosenburg, and M.B. Dickinson. The effects of prescribed burning on the thermal landscape. World Congress of Herpetology. 8-14 August 2012. (Poster Presentation)
- Howey, C.A.F., W.M. Roosenburg, and M.B. Dickinson. The effects of prescribed burning on the black kingsnake: going beyond the demographic data. Joint Meeting of Ichthyologists and Herpetologists. 6-11 July 2011. (Poster Presentation)
- Howey, C.A.F., W.M. Roosenburg, and M.B. Dickinson. The effects of prescribed burning on the black kingsnake: going beyond the demographic data. 4th Annual Fire in Eastern Oak Forests Conference. 17-19 May 2011. (Poster Presentation)
- Howey, C.A.F., W.M. Roosenburg, and M.B. Dickinson. The energetic expenditures of the black kingsnake (Lampropeltis nigra) in habitat altered by frequent prescribed burning. Joint Meeting of Ichthyologists and Herpetologists. 7-12 July 2010. (Poster Presentation)
- Howey, C.A.F., and S.A. Dinkelacker. Population demographics of the alligator snapping turtle (Macrochelys temminckii) in Arkansas. Annual Symposium on Conservation and Biology of Freshwater Turtles and Tortoises. 5-8 August 2009. (Oral Presentation)
- Howey, C.A.F., and S.A. Dinkelacker. Habitat selection of the alligator snapping turtle (Macrochelys temminckii): correlates between sex and reproduction. Annual Symposium on Conservation and Biology of Freshwater Turtles and Tortoises. 25-28 July 2007. (Oral Presentation)
- Howey, C.A.F., and S.A. Dinkelacker. Population demographics of the alligator snapping turtle (Macrochelys temminckii): evidence for past exploitation and present recovery. Annual Symposium on Conservation and Biology of Freshwater Turtles and Tortoises. 25-28 July 2007. (Poster Presentation)
- Howey, C.A.F., and S.A. Dinkelacker. Population demographics of the alligator snapping turtle (Macrochelys temminckii): evidence for past exploitation and present recovery. Annual Meeting of the Arkansas Academy of Science. 13-14 April 2007. (Oral Presentation)
- Howey, C.A.F., and S.A. Dinkelacker. Population demographics of the alligator snapping turtle (Macrochelys temminckii): evidence for past exploitation and present recovery. Southeastern Working Group of Partners in Amphibians and Reptile Conservation Annual Meeting. 22-25 February 2007. (Poster Presentation)
- Participated in “Future Collaborative Research and Conservation Goals for the Alligator Snapping Turtle” panel discussion, Moderator: Pritchard, P.C. Annual symposium on Conservation and Biology of Freshwater Turtles and Tortoises. 10-13 August 2006.
- Howey, C.A.F., and S.A. Dinkelacker. Thermoregulation in alligator snapping turtles (Macrochelys temminckii): correlates of habitat use, sex, and reproduction.Annual symposium on Conservation and Biology of Freshwater Turtles and Tortoises. 10-13 August 2006. (Poster Presentation)
- Howey, C.A.F., and S.A. Dinkelacker. Population demographics of the alligator snapping turtle (Macrochelys temminckii): evidence for past over-exploitation and present recovery. Joint Meeting of the American Society of Ichthyologists and Herpetologists. 12-17 July 2006. (Poster Presentation)
- Howey, C.A.F., and S.A. Dinkelacker. Population demographics of the alligator snapping turtle (Macrochelys temminckii): evidence for past over-exploitation and present recovery. Annual Meeting of the Arkansas Academy of Science. 7-8 April 2006. (Poster Presentation)

